Phylogeny
is evolutionary history of species or group of related species; provides important information on similar characteristics in closely related species; was used to identify species of whale from which 'whale meat' originated; bracketing allows people to predict features of ancestor from features of its descendants; analysis shows HIV is descended from viruses that infect chimpanzees and other primates.
Systematics
is discipline that classifies organisms and determines their evolutionary relationships.
Taxonomy
is scientific discipline concerned with classifying and naming organisms.
Carlos Linnaeus
published system of taxonomy based on resemblances in 18th century.
Key Features in Linnaeus' system
include two part names for species and hierarchical classification that remain useful today.
Binomal
is two part species scientific name.
Genus
is first part of name.
Specific Epithet
is second part that is unique for each species within genus.
Hierarchical Classification
is system Linnaeus introduced for grouping species in higher inclusive categories.
Taxonomic Groups
from broad to narrow include: 1)domain 2)kingdom 3)phylum 4)class 5)order 6)family 7)genus 8)species
Taxon
is taxonomic unit at any level of hierarchy.
Phylogenetic Tree
is evolutionary history of group of organisms; represents hypothesis about evolutionary relationships; for best hypothesis fit most data (morphological, molecular, and fossil).
Linnaean Classification
and phylogeny can differ from each other.
Systematists
have proposed classification system that would recognize only groups that include common ancestors and all its descendants; get information about morphologies, genes, biochemistry of living organisms; need to distinguish whether similarity is homology or analogy when making phylogeny; use computer programs, math tools when analyzing comparable DNA segments from different organisms; can never be sure of finding best tree in large data sheet; narrow options by applying max parismony and max likelihood.
Branch Point
represents divergence of 2 species.
Sister Taxa
are groups that share immediate common ancestor.
Rooted Tree
includes branch to represent last common ancestor of all taxa in tree.
Basal Taxon
diverges early in history of group and originates near common ancestor of group.
Polytomy
is branch which more than two groups emerge.
Polygenetic Trees
show patterns of descent (not phenotypic similarity); do not indicate when species evolved or how much change occurred in lineage.
Phylogenies
are inferred from morphological and molecular data.
Homology
is similarity due to shared ancestry (Ex: bat and bird wings [fare limbs]).
Analogy
is similarity due to convergent evolution. (Ex: bats and birds [functions of wings])
Homoplasies
are analogous structures or molecular sequences that evolved independently; molecules are identified by math tools.
Homologous
elements are more likely when they are similar in two complex structures; characters can be used to infer phylogeny once they have been identified.
Molecular Similarities
is important when distinguishing homology from analogy.
Shared Characters
are used to construct phylogenetic trees; along with different characteristics are in comparison for an organism with its ancestor; can be both ancestral and derived (depending on context).
Cladistics
group organisms by common descent.
Clade
is species group that includes ancestral species and all its descendants; a shared character first appeared in is useful to know when inferring to evolutionary relationships.
Monophyletic
is valid clade signifying that is has ancestor species and all its descendants.
Paraphyletic
is grouping that consists of ancestral species and some descendants.
Polyphyletic
is grouping that has distantly related species but does not include their most recent common ancestor; groups are distinguished from paraphyletic groups since they do not include most recent common ancestor.
Shared Ancestral Character
is character that originated in an ancestor of taxon; are characters shared by out-group and in-group that predate divergence of both groups from common ancestor.
Shared Derived Character
is evolutionary novelty unique to particular clade.
Out-group
is species or group of species that is closely related to in-group; is group diverged before in-group; are used by systematics to differentiate between shared derived, and shared ancestral characteristics.
In-group
is various species being studied.
Branch Length
can reflect on number of genetic changes that have been taken place in certain DNA sequence in that lineage; can represent chronological time, can be determined from fossil record.
Max Parismony
assumes that tree that requires fewest evolutionary events is most likely.
Max Likelihood
is principle that states that tree can be found that reflects most likely sequence of evolutionary events given certain rules about how DNA changes overtime.
Genome
is where an organism's evolutionary history is documented
Branch Points
from hundreds of millions of years ago can be useful when DNA that codes for r-RNA changes relatively slowly.
mt-DNA
evolves rapidly and can be used to explore recent evolutionary events.
Gene Duplication
increases number of genes in genome (providing more opportunities for evolutionary changes); result in gene families; can be traced to common ancestor like homologous genes.
Orthologous Genes
is homology that results from speciation event and occurs between genes found in different species; are widespread and extend across many widely varied species (Ex: humans, mice diverged about 65 million years ago, 99% of human genes are orthologous); has nucleotides substitutions that are assumed to be proportional to time since they last shared common ancestor.
Speciation Product
are orthologous genes.
Paralogous Genes
result from gene duplication (multiple copies of these genes; can diverge within clade that carries them and often evolve new functions; has nucleotides substitutions that are proportional to time since genes became duplicated.
Molecular Clocks
help track evolutionary time; uses constant rates of evolution in some genes to estimate absolute time of evolutionary change; are calibrated against branches whose dates are known from fossil record; does not run as smoothly as expected if mutations were neutral; application of HIV suggests that that strain spread to humans during 1930s; more advanced approach estimated first spread to humans around 1910.
Tree of Life
understanding continues to change based on new data; branches have had insight from people through molecular systematics lately; suggests eukaryotes and archaea are more closely related to each other than to bacteria; is based largely on r-RNA genes (some other genes reveal different relationships).
Kingdoms
Monera (prokaryotes), protista, plantae, fungi, animalia
Domain System
Bacteria, archaea, eukarya
Horizontal Gene Transfer
is movement from one genome to another; occurs by exchange of transposable elements, plasmids, viral infection, and organisms fusion; occurrence is result of disparities between gene trees; has played key role in evolution of both prokaryotes and eukaryotes; is argued by some biologists to be so common that early history of life should be represented as tangled network of connected branches.